Problems of Growth
Nineteenth Ischia Summer School on the History of the Life Sciences
Biblioteca Antoniana, Ischia, Italy, 28 June – 5 July 2026
Applications are invited for this week-long summer school, which provides advanced training in history of the life sciences through lectures, seminars and discussions in a historically rich and naturally beautiful setting. The theme for 2026 is 'Problems of Growth’. The deadline is Friday 27 February 2026.
Organizers: Christiane Groeben (Naples, local organizer), Nick Hopwood (Cambridge), Erika L. Milam (Princeton), Staffan Müller-Wille (Cambridge) and the Stazione Zoologica Anton Dohrn
Confirmed faculty: Daryn Lehoux (Queen’s, Canada), Dániel Margócsy (Cambridge), He Bian (Princeton), Patrick Anthony (Uppsala), Alison Bashford (UNSW), Hannah Landecker (UCLA), Edna Suárez-Díaz (UNAM), Sabina Leonelli (TU München)
For funding we are most grateful to Cambridge HPS, Cambridge Intesa Sanpaolo Fund, George Loudon, Stazione Zoologica Anton Dohrn, Dohrn Foundation, Science History Institute, Centro Etnografico delle Isole Campane, Center on Science and Technology at Princeton University and the Italian Society for the History of Science.
More information: <http://ischiasummerschool.org/>
About the school
The Ischia Summer School on the History of the Life Sciences provides advanced training in a lively international field that offers a long-term perspective on some of the most significant ideas, practices and institutions in the world today. The school, which has a tradition of association with the Naples Zoological Station, was revived in 2005 after a break of two decades and has run every other year since then other than during the coronavirus pandemic. We can accommodate up to 26 graduate students and postdoctoral fellows. The event provides a structured learning experience plus extensive opportunities for participation and interaction. English is the working language and we encourage exchange of ideas across disciplinary boundaries, national cultures and historical periods. Spending the week on an island, staying in the same hotel and sharing breaks and meals maximizes opportunities for exchange. These are enhanced through social events, including a welcome reception and a day trip to Naples, the morning spent learning about the history and current research of the Station, the afternoon free for sightseeing. There will also be a free afternoon to explore Ischia itself.
Introduction to the theme
Growth affords hope and attracts fear. Balanced growth feeds populations, fuels prosperity and imparts purpose to individual and collective lives. The unfettered growth of cells, pathogens, parasites and populations threatens physiological, economic and ecological collapse. Even balance may be a problematic ideal: norms of flourishing and beauty have guided discrimination by vaunting harmonious over retarded, excessive or monstrous growth. The sustainability of life on Earth, attempts ‘to change the story of cancer’ and the politics of human diversity: growth is at the heart of them all. Yet compared with other vital processes, notably inheritance, development and reproduction, growth in the life sciences has lacked status and attention. This summer school provides an opportunity to explore knowledges and practices of growth between antiquity and the present day while bringing together problems usually kept apart.
For Aristotle, vegetative growth was the lowest function of the soul and for that reason fundamental to plants, beasts and humans. Unlike fire, vegetative growth had a natural limit. Where minerals grew by external accretion or juxtaposition, living beings had the distinctive ability to expand by assimilation of nutrients from the inside out, whether organ by organ or from a preformed seed. Surgeons tried to remove those tumours, cankers and warts that resulted from an imbalance of humours among other causes. Generation, which was hard to imagine in mechanical terms, was often framed as a special form of growth. Late medieval philosophers brought together generation, projectile movement and the accumulation of capital as sharing the same basic problem, how a movement severed from its mover could continue to produce. In a balanced world, gain in one part was compensated by loss elsewhere. Large animals, according to Aristotle, produced fewer offspring, and the relative growth of one organ entailed the diminution of another. At Italian universities during the Renaissance, these ancient ideas were taken up and reformed by scholars including Girolamo Fabrici d’Acquapendente, Andrea Cesalpino and Marcello Malpighi in attempts to reground the systematic study of nature and naturalize growth and development.
By contrast, it seems, modern approaches to growth, in biology as in economics, aimed for an overall increase—in size, in number of individuals and in productivity. As the ultimate source of economic progress the physiocrats postulated an inherent capacity of nature to reproduce. Naturalists like Lazzaro Spallanzani located the same reproductive and regenerative capacities in minute parts that made up animal bodies. But proper growth was also reckoned to occur within certain limits. In the principle of population Thomas Robert Malthus expressed the limit set for the potentially geometric growth of human numbers by the merely arithmetic growth of food supplied from the land. More generally, in the hands of the population biologist Raymond Pearl the S-shaped curve came to capture the colonization of a new space, with slow initial acceleration towards exponential growth and then deceleration as environmental resistance increased and the ‘carrying capacity’ was reached. Based on computer simulations of the catastrophic consequences of runaway population and economic growth, the Club of Rome’s bestselling report The Limits to Growth (1972) is a point of origin for debate over ‘degrowth’ and ‘sustainable growth’.
Classical discussion of growth within organisms had been informed by the canons of beauty appropriate to each stage of life, with more attention to proportion than size. Beginning in the eighteenth century, longitudinal measurements of human growth aligned with demands for military manpower and projects of social reform. Measurement fed debate over the roles of heredity and environment. On the one hand, anthropometry ultimately produced distinct growth equations for groups defined by age, sex and race. Unbalanced growth was associated with monstrosity and other ways of falling short of the white, male model. On the other, failure to grow became an index of deprivation, most obviously, as physiologist Angelo Mosso argued, in the stunting of factory children. Eugenicists, notably criminologist Cesare Lombroso, were concerned with imbalance at the level of populations.
Standards justified clinical intervention in pathologies of growth. James Tanner, who led the Harpenden study into growth through puberty into adulthood, pioneered the treatment with growth hormone of children who looked set to miss out on the advantages of height. Since the 1980s ultrasound measurements of fetuses have identified growth restrictions on an ever larger scale. Yet even after major surveys from Turin to Nairobi, it is controversial to what extent the standards should be universal or tailored to demographic groups.
In the nineteenth century the knotty issues involved in defining individuals that were explored productively at the Stazione Zoologica di Napoli made growth hard to distinguish from maintenance and reproduction. An influential formulation held that reproduction represented growth beyond the individual limit. From the 1860s embryonic development was discussed in terms of the differential growth of parts. Inspired by D’Arcy Thompson’s On Growth and Form (1917), Julian Huxley set an agenda with Problems of Relative Growth (1932) and the notion of allometry, or the shape-changing growth of a part at a different rate from the organism as a whole. Mechanisms could be studied in ontogeny or changing patterns traced in phylogeny. In a famous essay, ‘On being the right size’, J.B.S. Haldane proposed that ‘Comparative anatomy is largely the story of the struggle to increase surface in proportion to volume’: more complicated forms enable the larger sizes that maintain body temperature at lower metabolic rates.
Within a species, tissues and organs must somehow ‘know’ when to stop growing. The cell theory framed organismal growth as the division and expansion of these elementary parts. Cancer, the disease that made biomedicine, came to be understood as a pathology of malignant growth. Research elucidated factors, not least growth factors, notably nerve growth factor discovered by Stanley Cohen and Rita Levi-Montalcini, that promoted, regulated and interfered with cell division. Alongside chemotherapies, weedkillers were developed that acted by causing rapid, uncontrolled growth. Synthetic auxins, the hormones that regulate cell division and expansion in plants, became notorious as the defoliant Agent Orange used by the British in the Malayan Emergency and the United States in the Vietnam War.
This sketch raises large questions. Should understandings and practices of growth be seen as having first sought balance, then promoted unlimited increase before recognition of the costs of growth called the whole framework into question? Or did gospels of growth acknowledge the need for some balance? Should we grasp growth as a modern or capitalist imperative, a potentially relentless power and a creative one through the transformation of quantity into quality? Or is a reason for its neglect in reflection on the life sciences (as distinct from economics and agronomy) that growth implies mere increase in size or number while the truly remarkable changes have seemed to result from qualitative alterations? Reflexively, reservations about growth apply to knowledge, too; simply accumulating data has seemed inadequate when we might need a whole new paradigm. A long-term theme and implicated in urgent problems, growth in and around the life sciences provides a rich field for historical deliberation and for trade between disciplines.
Programme
The school starts with registration and a reception on the afternoon of Sunday 28 June, and ends after dinner the following Saturday night. Departure is on Sunday 5 July. Lectures last for up to 30 minutes in one-hour slots, leaving at least 30 minutes for discussion. Seminars focus on pre-circulated texts. Groups of students will prepare each one with the seminar leader.
Daryn Lehoux (Queen’s, Canada)
Lecture: Aristotle on nutrition, growth, residues and seed
Seminar: The ‘faculty’ of growth in Galen
Dániel Margócsy (Cambridge)
Lecture: Soil, vermin and ghosts: The limits to growth in agriculture and medicine in early modern Europe and Indonesia
Seminar: Humans and horses: Theorising size in early modern European Medicine
He Bian (Princeton)
Lecture: Growth and regeneration in early modern Chinese thought
Seminar: Growing empire, coining new names: Manchu as a language for flora and fauna nomenclature
Patrick Anthony (Uppsala)
Lecture: Toward a history of extractive sciences—and the end of the mineral frontier
Seminar: From bio-geography to necro-geography: Sciences of life and death during the Circassian genocide
Alison Bashford (UNSW)
Lecture: Growth, limits and the afterlife of Malthus
Seminar: Fertility decline and modernity’s great deceleration: Where is reproduction/population in degrowth scholarship?
Hannah Landecker (UCLA)
Lecture: The butcher’s philosophy: Transmuting knowledge of life into knowledge of growth in modern agriculture and medicine
Seminar: Practical approaches to working with visual documents: Exploring cases and patterns in an industrial trade journal archive
Edna Suárez-Díaz (UNAM)
Lecture: Geographies of malnutrition: The clinic, the lab and the committee
Seminar: Traditions of knowledge and intervention: Studying malnutrition and mental development in the land of Zapata
Sabina Leonelli (TU München)
Lecture: Growing data crops: Extractivism and agriculture
Seminar: Colonial trends in agricultural data sharing
Public lecture: Intelligenza ambientale: Come usarla per salvare il pianeta
Cost
The fee for students is €400 each, which includes hotel accommodation and all meals for the week. Students need to pay for their own travel to Ischia. The directors will consider requests to waive the fee for accepted students unable to raise the money themselves, when supported by a detailed financial statement and a letter from their department head.
Applications
Applications should be sent by email to <administrator@ischiasummerschool.org> and should include, please:
• a statement specifying academic experience and interest in the course topic (max. 300 words),
• a brief CV,
• a letter of recommendation.
The deadline for applications is midnight CET on Friday 27 February and applicants will be notified of the outcome by 13 March 2026.
